Most lionesses will have reproduced by the time they are four years of age.Lions do not mate at any specific time of year, and the females are polyestrous.As with other cats' penises, the male lion's penis has spines that point backward. During withdrawal of the penis, the spines rake the walls of the female's vagina, which may cause ovulation.A lioness may mate with more than one male when she is in heat; during a mating
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